.

Wang, Z., Skidmore, A. K., Wang, T., Darvishzadeh, R., & Hearne, J. (2015). Applicability of the PROSPECT model for estimating protein and cellulose+ lignin in fresh leaves.  Remote sensing of environment ,  168 , 205- Physical models also include empirical parts. PROSPECT does so, too. In the paper, leaf potein and lignin + cellulose were decoupled from LMA.

Li, F., et al. (2017). "Foliar nitrogen and phosphorus stoichiometry of three wetland plants distributed along an elevation gradient in Dongting Lake, China." Sci Rep 7(1): 2820. Species specific elemental ratio constraint its distribution partially. Then plants spatial distribution can be explained by their stoichiometry.