Stenberg, P. (1998). Implications of shoot structure on the rate of photosynthesis at different levels in a coniferous canopy using a model incorporating grouping and penumbra. Functional Ecology, 12(1), 82-91.
In the case of conifer, needles are too small to make umbra. Photosynthesis would mainly occur in penumbra, which is outside of direct sunlight. Conifer increases their LAI by grouping their leaves. It can result the decrease in total light capturing. However, the amount of photosynthesis increasing in lower leaves, leaves can more efficiently capture the light.
In the case of conifer, needles are too small to make umbra. Photosynthesis would mainly occur in penumbra, which is outside of direct sunlight. Conifer increases their LAI by grouping their leaves. It can result the decrease in total light capturing. However, the amount of photosynthesis increasing in lower leaves, leaves can more efficiently capture the light.
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